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indignant flight

Hi,

I would thankful if you could say whether indignant flight here means those crows flied angrily, since they had to fly backward?

Out of the sewage Biju chased a line of pigs with black watermarks across their bellies, ran after them in jubilation. "Hup hup," he shouted. The crows that had been sitting on the pig's backs scrambled into indignant flight, having to start up backward.

Many thanks.
Chapter 30 of Inheritance of Loss, a novel by Kiran Desai.
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