| Foskeia | |
|---|---|
| 👁 Image | |
| Snout elements of the paratype (A) and holotype (B-C) of F. pelendonum | |
| Scientific classification 👁 Edit this classification | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Clade: | Dinosauria |
| Clade: | †Ornithischia |
| Clade: | †Ornithopoda |
| Clade: | †Rhabdodontomorpha |
| Genus: | † Dieudonné et al., 2026 |
| Species: | †F. pelendonum
|
| Binomial name | |
| †Foskeia pelendonum Dieudonné et al., 2026
| |
Foskeia is an extinct genus of rhabdodontomorph ornithopod dinosaurs known from the Early Cretaceous (Barremian to Aptian ages) Castrillo de la Reina Formation of Spain. The genus contains a single species, Foskeia pelendonum, known from fossils of multiple individuals of different ontogenetic stages. Before its formal description, it had been nicknamed the "Vegagete ornithopod."[1]
History of discovery
[edit]In 2016, Dieudonné et al. reported the discovery of at least five individuals of a gracile, small-bodied ornithopod (approximately 0.5 m (1.6 ft) long), representing several distinct ontogenetic stages. Despite the abundance of material, the describers refrained from assigning a binomial name to it, instead referring to it as the "Vegagete ornithopod," after the locality where the fossils were found. This locality is part of the Castrillo de la Reina Formation (Cameros Basin) in Burgos Province, Spain, a site known for its rich diversity of Early Cretaceous dinosaur remains.[2]
The discovery of new cranial elements in subsequent fieldwork and assessment of this material via μCT scanning allowed the remains to be formally described as a new genus and species of rhabdodontomorph ornithopod in 2026.[1] The generic name, Foskeia, is derived from the Ancient Greek words φῶς (light) and βόσκειν (to forage), referring to its small size and presumed herbivorous habits. The specific name, pelendonum, is a reference to the Pelendones, a Celtiberian tribe from the Fuentes de Duero, near the type locality.[3]
Paleobiology
[edit]Analysis of the limb bones performed by Dieudonné et al. (2023) revealed that Foskeia juveniles began life as quadrupedal animals. However, at an early stage in their ontogeny, they transitioned to bipedalism. Histological analysis of the limb bones further revealed that the largest known individual of Foskeia represents a late subadult, which supports the interpretation that the species was among the smallest ornithopods. The study showed that the ontogenetic pathway of Foskeia involved significant skeletal changes, with muscle attachment marks observed on the femora and tibiae indicating the gradual development of a more dynamic and efficient locomotor system as the animal matured. This postural transition is consistent with other early rhabdodontomorphs.[4]
Classification
[edit]When it was first described by Dieudonné and colleagues in 2016, the then-unnamed Vegagete ornithopod was found to be a basal member of the newly named clade Rhabdodontomorpha, diverging after Muttaburrasaurus as the sister-taxon of the Rhabdodontidae. A cladogram adapted from their analysis is shown below:[2]
| Iguanodontia |
|
In 2025, Czepiński & Madzia re-analyzed all rhabdodontomorph material known at that time. Scoring their results in the comprehensive ornithischian-focused phylogenetic analysis of Fonseca et al. (2024),[5] they recovered the Vegagete ornithopod as a member of the Rhabdodontoidea. The broader rhabdodontomorph clade was recovered in a basal (primitive) position within the Iguanodontia. The rhabdodontomorph results of their analysis are shown below:[6]
| Rhabdodontomorpha |
|
In 2026, upon the formal naming of the Vegagete ornithopod, Dieudonné et al. placed the taxon within a novel subclade within the Rhabdodontomorpha, which they named Rhabdodontia. Foskeia was recovered diverging after Muttaburrasaurus and Fostoria, as the sister taxon to a traditional Rhabdodontidae. Contrary to previous analyses, the rhabdodontomorph clade was placed in a more derived (advanced) position, at the base of the Ankylopollexia. A cladogram adapted from their analysis is shown below:[1]
| Iguanodontia |
|
References
[edit]- ^ a b c Dieudonné, Paul‐Emile; Becerra, Marcos Gabriel; Zanesco, Tábata; Tortosa, Thierry; Cruzado‐Caballero, Penélope; Stein, Koen; Torcida Fernández‐Baldor, Fidel (2026-02-01). "Foskeia pelendonum, a new rhabdodontomorph from the Lower Cretaceous of Salas de los Infantes (Burgos Province, Spain), and a new phylogeny of ornithischian dinosaurs". Papers in Palaeontology. 12 (1). doi:10.1002/spp2.70057. ISSN 2056-2799.
- ^ a b Dieudonné, Paul-Emile; Tortosa, Thierry; Fernández-Baldor, Fidel Torcida; Canudo, José Ignacio; Díaz-Martínez, Ignacio (2016). "An Unexpected Early Rhabdodontid from Europe (Lower Cretaceous of Salas de los Infantes, Burgos Province, Spain) and a Re-Examination of Basal Iguanodontian Relationships". PLOS ONE. 11 (6) e0156251. Bibcode:2016PLoSO..1156251D. doi:10.1371/journal.pone.0156251. PMC 4917257. PMID 27333279.
- ^ Free University of Brussels. "Tiny new dinosaur Foskeia pelendonum fills in an evolutionary gap". Phys.org. Retrieved 2026-02-02.
- ^ Dieudonné PE, Torcida Fernández-Baldor F, Stein K (2022). "Histogenesis and growth dynamics of the tiny Vegagete rhabdodontomorph hindlimb (Ornithischia, Ornithopoda): paleoecological and evolutionary implications". Cretaceous Research. 141 105342. doi:10.1016/j.cretres.2022.105342. S2CID 251984848.
- ^ Fonseca, André O.; Reid, Iain J.; Venner, Alexander; Duncan, Ruairidh J.; Garcia, Mauricio S.; Müller, Rodrigo T. (2024). "A comprehensive phylogenetic analysis on early ornithischian evolution". Journal of Systematic Palaeontology. 22 (1) 2346577. Bibcode:2024JSPal..2246577F. doi:10.1080/14772019.2024.2346577.
- ^ Czepiński, Łukasz; Madzia, Daniel (2025-04-30). "Exploring the diversity and disparity of rhabdodontomorph ornithopods from the Late Cretaceous European archipelago". Scientific Reports. 15 (1): 15209. Bibcode:2025NatSR..1515209C. doi:10.1038/s41598-025-98083-z. ISSN 2045-2322. PMC 12044058. PMID 40307357.
- ^ Maidment, Susannah C. R.; Butler, Richard J.; Brusatte, Stephen L.; Meade, Luke E.; Augustin, Felix J.; Csiki-Sava, Zoltán; Ősi, Attila (2026-01-07). "A hidden diversity of ceratopsian dinosaurs in Late Cretaceous Europe". Nature: 1–7. doi:10.1038/s41586-025-09897-w. ISSN 1476-4687.
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